>>4609243
They were grouped up more for sure, but the puzzles were much more complex and varied. In the end, Exoddus had way more content. It was also a little gimmicky in the first game to have puzzles where you had to complete it in multiple passes because only one Mudokon could be commanded at a time.
>>4609509
Strange design choice, especially for how early in the game they are encountered. I imagine it was a stumbling block for a lot of players, especially those that didn't play the first game. Still the most stressful part of either game for me, even though when you're familiar with the game they are trivial to deal with.